Spinning slowly in circles in his small concrete tank in an off-limits area of an aquarium, Romeo the manatee is the picture of loneliness.
The 67-year-old has been at the Miami Seaquarium - the same center where the 'loneliest whale in the world', Lolita, died earlier this year - since 1956, and campaigners say he has been kept in isolation since at least April.
Manatees are semi-social animals and 'suffer psychologically when not living in pairs or groups' - but Romeo has long been separated from his partner, and the mother of his offspring, Juliet, and has been alone since his other companions were released.
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